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Seems to spawn at random. Found mine on a barroth investigation in the desert area with the sand pit trap.
Elusive as *****
keep reloading the area and head to the highest point of the map he schould be in the nest expeditions work
Start in wildspire waste area 6, after exiting the camp go left, climb up and check the cactus field on the left side, then jump down and check the cactus field on the other side. then walk a few steps towards the blissbill nest and use binuculars to check if there is a cactuar jumping on top of it. Travel to another area and back to wildspire waste and repeat.