Dedicated to digging into the game's lore. Bring your thinking caps.
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cyseal

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Battle Tested

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#1
From what creature is the skull in ash lake?
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CakeThiefPro

Obsessed
Commander

Joined: Thu Mar 28, 2013 9:00 pm
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#2
It's one of those mysteries that has no real answer. The most likely answer is that it's from some kind of dragon considering the location or perhaps it belongs to a more powerful form of giant that was wiped out amidst the war with the dragons. Sadly it's never been referenced again and there's no strong theory on the skull.
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Dapiek_Absaroka

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First Warden

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#3
I always found it too humanoid to consider it to be from a dragon, so my bet would be that it's from a giant. Granted, it would be from a particularly large and bestial variety of giant but considering how varied in size and morphology giants are in the Dark Souls games it's not exactly outside the realm of possibility.